

The analytical form for RED circumnavigates the need to spin up the numerical model, making it attractive for application in Earth system models (ESMs). Using net primary productivity and litter outputs from the UK Earth System Model (UKESM), we are able to diagnose the spatially varying disturbance rates consistent with this observed vegetation map. We show that RED can thus be calibrated to the ESA LC_CCI (European Space Agency Land Cover Climate Change Initiative) coverage dataset for nine PFTs. The analytical equilibrium solutions of RED allow the model to be calibrated against observed forest cover using a single parameter – the ratio of mortality to growth for a tree of a reference mass ( μ 0). The population of each plant functional type (PFT) is partitioned into mass classes with a fixed baseline mortality along with an assumed power-law scaling of growth rate with mass. RED is designed to retain the ability of more complex cohort DGVMs to represent forest demography, while also being parameter sparse and analytically solvable for the steady state. Here we present the Robust Ecosystem Demography (RED), in which the pdf is collapsed onto the single dimension of tree mass. This approach can capture the overall impact of stochastic disturbance events on the forest structure and biomass – but at the cost of increasing the number of parameters and ambiguity when updating the probability density function (pdf) in two dimensions.

These models typically attempt to simulate the size distribution of trees as a function of both tree size (mass or trunk diameter) and age (time since disturbance). More advanced cohort-based patch models are now becoming established in the latest DGVMs. Dynamic global vegetation models (DGVMs) vary in their representations of regrowth and competition for resources, which results in differing responses to changes in atmospheric CO 2 and climate. A significant proportion of the uncertainty in climate projections arises from uncertainty in the representation of land carbon uptake.
